Species Synonyms
Within botanical treatises and collection registers, an extensive Latin nomenclature is employed to designate this species: Vitis flexuosa Thunberg, Vitis indica Thunberg, Vitis parvifolia Roxburgh, Vitis purani Don, Vitis wallichii de Candolle, and Vitis vulpina var. parvifolia Regel. Historical accounts also list Vitis truncata Blume, Vitis papillosa Herb. Delile, Vitis sylvestris Blume, and Vitis succisa Hance. Furthermore, several distinct varieties are recognized within its taxonomy: var. blumei Planchon, var. malaiana Miquel, var. thomsoniana Planchon, var. parvifolia Planchon, and var. gaudichaudii Planchon. In certain ancient Indian collections, the vine was occasionally misidentified under the name Vitis vinifera Hook. et Thoms.
Historical Background and Geographical Distribution
The scientific description and systematic classification of Vitis flexuosa were established by the viticulturist J. E. Planchon based on the examination of extensive herbarium collections. Due to significant external diversity and the variability of individual traits (polymorphism), researchers long questioned the specific identity of all its geographical forms, given that wild grape species generally occupy more localized ranges.
Vitis flexuosa exhibits an exceptionally broad geographical distribution across the Asian continent. Native populations of the species are documented in Japan, Korea, and various provinces of China (specifically in Yunnan). A distinct form, Malaiana, is distributed on the island of Java. In the southern regions of China and Cochin-China (modern-day Vietnam), the form Thomsoniana is prevalent. The form Parvifolia is recorded in Kashmir, and the vine successfully vegetates in the high-altitude regions of the Himalayas at elevations ranging from 1,500 to 1,800 meters above sea level (form Thomsoniana). In European scientific practice, it is specifically the Japanese form of the species that has been described in detail. The plant holds exclusively botanical interest, was never cultivated on a commercial scale in Europe, and stands clearly distinct from the species Vitis thunbergii and Vitis lanata, with which it was frequently confused in earlier periods.
Botanical and Morphological Description
- Growth Habit and Trunk: The vine is characterized by moderate vigor, with weakly pronounced branching. In its native habitat, it develops as a slender climbing vine. The older bark exfoliates in narrow longitudinal ribbons of a reddish-gray shade. The anatomical structure of the seeds is illustrated in figures (Fig. 716–717).
- Annual and Herbaceous Shoots: Developed canes are long, slender, straight, and cylindrical in cross-section. The nodes are very weakly defined and practically unexpanded. The internodes are long, with fine, barely perceptible longitudinal striations. Mature annual canes acquire a characteristic brownish-gray coloration. The surface of the cane is uniformly covered with short, soft, grayish hairs, resembling the pubescence seen in the tomentose forms of the American species Vitis riparia. The tendrils are intermittent, exceptionally long, slender, and bifurcated (bifid).
- Leaf: The leaves are small and entire, showing no traces of lateral sinuses or lobes. The margins of the leaf blade are regular and even. In shape, the leaf is elongated ovoid-elliptic, with the blade length noticeably exceeding its width. The petiolar sinus is shallow, widely open, and V-shaped. In the specific form Parvifolia, the leaf base can assume a hastate shape with a completely absent sinus base. The marginal teeth are weakly pronounced, and their positions along the smooth edge of the blade are indicated solely by short mucros. On the upper surface, the anatomical veins appear sunken into the tissue. On the lower surface, the veins are robust, quite prominent, and densely covered with short, soft hairs.
- Inflorescences and Berries: The inflorescences display a regular cylindrical shape, are elongated, and feature weak lateral branching. In their herbaceous state, the inflorescence axes and buds are completely embedded in a dense, whitish fluff. The peduncle is exceptionally long and slender. The clusters are small and loose. The berries are small and spherical, turning a matte black color upon full maturity. The pulp is lacking juiciness, and the juice is released in small quantities. The flavor is acidic and astringent.
- Seeds: Each berry develops between 2 and 4 small seeds. The seeds are round-trigonal, with an exceptionally short and thin beak. The coloration of the seed coat is glossy and of a dark brown or chestnut hue. The chalaza is oval, weakly prominent, and situated in the upper third of the dorsal side of the seed. The raphe is thin and thread-like.